> All kinds of animals would benefit from knowing that awhoo comes from a wolf and that, in this example, awhoo awhoo is the same sound coming from the same wolf or that an animal recognizes that the first awhoo comes from one wolf and the second awhoo from another wolf.

Interesting choice of example. Dog howls are apparently intentionally off-key.

The goal of the basic choir is to sound like one voice in perfect harmony.

The goal of a pack howling over long distances is apparently to communicate that every member of the pack is present - so each voice is out of harmony with the rest.