Little bro read the paper title and no further.

That study found that when you test 14 monkeys alone in cages where they can’t actually move the toys, you don’t see the same sex differences as when 135 monkeys are tested in social groups with freely movable toys.

The authors themselves say the social context may be necessary for expression. That’s not evidence against biological contribution, but evidence that behavior requires context to manifest.

You don’t disprove hunger by noting that people don’t eat when there’s no food available.