Every single one of those demons is also a chimera and so if you look, there’s a half elephant, half fish in the top left corner. If you look in the top right, there’s like a half fish half horse. So if you look at the gaping demon butthole again, and look at the rest of the features around the gaping opening. You’ll realize that it is the lower half of the creature and it is a sea creature you can see there’s like a coral tentacles thing behind it. It’s just a fish‘s mouth. That does also happen to function as the demon‘s butthole, apparently
This insight won me over on the butthole.