The open and closed apple keys first appeared on the Apple ///, initially next to each other on the left of the spacebar. On the Apple /// plus, the closed apple then moved to the right of the keyboard, which is what the Apple IIe inherited.

The closed apple key then appeared on the Lisa keyboard alongside an option key (both on the left of the spacebar), but the Lisa's closed Apple key acted like and is what became the Mac's command key.

https://www.nightfallcrew.com/09/12/2014/apple-iii-apple/

https://mirrors.apple2.org.za/Apple%20II%20Documentation%20P...

https://vintagecomputer.net/apple/lisa/apple_lisa_A6S0200_ke...