Tetris high scores, obviously

Mother nature hates weak things that die (that's why they get eliminated), so if we can make it to interplanetary species before killing ourselves, that would be a pretty huge sign of success. At least on mother natures benchmark.

Some of us don't spend days looking for food, don't die of cold, and survive the flu...

aaand we have Quake and Comand&Conquer - Red Alert

The most successful at communicating their view that they are the most successful. Whether they are or not. But that means they are. By that metric.

Has another animal proposed they are more successful by a different metric?

Crickets?