If you really think about why MoE came into existence, its to save significant cost during training, I don't think there was any concrete evidence of performance gains for comparable MoE vs dense models. Over the years, I believe all the new techniques being employed in post training have made the models better.
MoE models will have far more world knowledge than dense models with the same amount of active parameters. MoE is a no-brainer if your inference setup is ultimately limited by compute or memory throughput - not total memory footprint - or alternately if it has fast, high-bandwidth access to lower-tier storage to fetch cold model weights from on demand.
I think you mean inference compute? I believe all expert weights are updated in each backward pass during MoE training. The first benefit was getting a sort of structured pruning of weights through the mechanism of expert selection so that the model didn’t need to go through ‘unnecessary’ parts of the model for a given token. This then let inference use memory more efficiently in memory constrained environments, where non-hot or less common experts could be put into slow RAM, or sometimes even streamed off storage.
But I don’t think it necessarily saved training cost; if it did, I’d be interested to learn how!